Prioritize an evolution that fixes the current team blocker: Shadow, Puppet, or Cursed Student for a main carry; Salmon Sorcerer for control; Elf Mage for a Magical damage route. Search the recipe finder before converting Sprites or spending Gold, then open the live Evolve source panel for special items.
Unlocks the Arcane charge cycle and stronger Old Magic, Eruption, and Storm follow-ups.
If you are missing materials
Evolve before spending heavily on equipment rerolls, because Elven Battle Staff is built around the evolved spell cycle.
high
Flame Emperor → Flame Emperor (Reincarnate)
Last checked 2026-07-22
Required materials
1× Burning FruitOpen this item inside the Evolve menu and follow the current Source panel before farming
20× Red SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
20× Yellow SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
1× Rainbow SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
5000× GoldStory clears, Challenges, quests, and account rewards
Required map / mode
Open Burning Fruit in the Evolve menu; farm the current Sprite routes shown in progression rewards.
Exclusive equipment
Emperor’s Attire — Rose Kingdom Mastery or Armory Forge crafting.
Evolution benefit
Adds a placeable Wall of Fire and improves the Burn-spread route used by Hollow.
If you are missing materials
Raise priority when you own Hollow or another unit that gains damage from Burn; otherwise finish the main carry first.
high
Crimson → Crimson (Brother)
Last checked 2026-07-22
Required materials
20× Red SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
20× Pink SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
1× Rainbow SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
5000× GoldStory clears, Challenges, quests, and account rewards
Required map / mode
Story, Challenges, Mastery, Expeditions, and Raids can provide Sprites; check the current reward panel before repeating a stage.
Exclusive equipment
No exclusive equipment route confirmed for this evolution
Evolution benefit
Adds Piercing Crimson and strengthens the mark, explosion, and damage-pool wave-clear loop.
If you are missing materials
Reserve the Pink and Red Sprites once Crimson is confirmed in your active team; the recipe has no separate special item at this check.
medium
String Demon → String Demon (Awakened)
Last checked 2026-07-22
Required materials
1× Joker of the Nonsensical PiratesOpen this item inside the Evolve menu and follow the current Source panel before farming
20× Yellow SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
20× Pink SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
1× Rainbow SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
5000× GoldStory clears, Challenges, quests, and account rewards
Required map / mode
Open the Joker item in the Evolve menu for the current source; collect Sprites from active progression modes.
Exclusive equipment
Webbed Fruit — Rose Kingdom Mastery or Armory Forge crafting.
Evolution benefit
Adds String Sync follow-up attacks and strengthens Cocoon-based control.
If you are missing materials
Evolve after the primary carry unless the current blocker is crowd control or follow-up support rather than raw damage.
bridge
True Saint → True Saint (Holy)
Last checked 2026-07-22
Required materials
1× Holy PendantOpen this item inside the Evolve menu and follow the current Source panel before farming
5× Red SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
5× Blue SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
5× Green SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
5× Purple SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
5× Yellow SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
5× Pink SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
1× Rainbow SpriteStory, Challenges, weekly Challenges, Mastery, Expeditions, or Raids; confirm the live reward panel
5000× GoldStory clears, Challenges, quests, and account rewards
Required map / mode
Check Holy Pendant in the Evolve source panel; the recipe uses a smaller mixed-color Sprite set.
Exclusive equipment
No exclusive equipment route confirmed for this evolution
Evolution benefit
Adds Seize, temporarily borrowing damage from the strongest nearby units.
If you are missing materials
Use as a progression bridge when the mixed Sprite cost is affordable, but avoid blocking a top carry evolution for it.
No matching recipes
Try the base unit, evolved form, a Sprite color, Gold, or the special item name.
Material routes
Farm the missing category
Colored Sprites
Check Story, Challenges, weekly Challenges, Mastery, Expeditions, and Raids. Use the current reward panel.
Rainbow Sprites
Keep flexible Sprite resources until one evolution is selected; higher-tier rewards or crafting may be required.
Special item
Select the item in the Evolve menu and follow its live Source panel. Availability can rotate.
Gold
Reserve the full recipe cost before spending on shop or equipment upgrades.
Before traits and equipment
Lock the evolved job first
1
Confirm the unit already solves a real Story, Raid, Challenge, or Expedition problem.
2
Preview the evolved passive and exact cost in the lobby Upgrades room.
3
Collect the special item before converting flexible Sprite stock.
4
Evolve, retest the unit, then choose the trait and exclusive equipment route.
Continue with this game
More Anime Expeditions pages
Open the next guide, database, update, or tool when the current decision is complete.
Player questions
Evolution FAQ
Use the current game panel before spending rare resources on a value that can change.
Where do Evolution Sprites come from?
Sprites can come from Story, Challenges, weekly Challenges, Mastery, Expeditions, and Raids. Check the current reward panel because the best repeat stage can change.
Where do I find a special evolution item?
Open the unit in the Evolve menu, select the missing item, and use its current Source panel. Special-item availability can rotate, so this is safer than following an old fixed map claim.
Which unit should I evolve first?
Choose the unit already carrying several modes. Shadow, Puppet, and Cursed Student are top damage priorities; Salmon Sorcerer can be first when control is the reason runs fail.
Should I roll traits before evolving?
Usually wait until the unit has earned a long-term slot. Evolution changes the kit and can change whether Unbound, Draconic, Forsaken, Primordial, or another trait is the best investment.
Can I preview an evolution before spending?
Yes. The Evolve menu shows requirements and the evolved passive. Compare that benefit with the material cost before pressing Evolve Unit.
